生活型

climber

形態特徵

• Stems 6-9 m long, 3-12 mm thick, subterete or obtusely 3- or 4-angular, climbing; internodes 7-30 cm long, 3-12 mm thick; branches 30-40 cm long, 2-3 mm thick, weakly zigzag, obtusely 3- or 4-angled, dull, striate, smoothish or sparingly prickly; internodes 1-6 cm long, 2-3 mm thick.
• Leaf blade broadly ellipsoid to broadly ovate-ellipsoid, rarely ovate-orbicular, 5-10(-13) cm long, 3-6(-10) cm wide, base rounded or suddenly contracted, apex subacute or suddenly contracted, mucronate, thick herbaceous to thin coriaceous; costas 7 including a fine marginal pair, 5 main costas rather sharply raised on lower surface, median 3 often united for 3-7 mm above base, lateral veinlets oblique, many, curved upward, some further divided, reticulations not elongated, relatively loose; petiole 8-20 mm long, 1.2-1.4 mm thick, weakly laterally compressed, with a sharp abaxial angle; sheath 5-6 mm long; wings 1- 1.5 mm wide, herbaceous to membranaceous, auricle deltoid, ca. 1 mm long; tendrils on both stem and branch leaves, 6-20 cm long.
• Inflorescence in prophyllate axils on lower part of branches, both sexes with 3-7 umbels; common axis 3-6 cm long, relatively straight, internodes 1-3 cm long; prophyll large, ovate, subacute, 7-9 mm long, 3-4 mm wide, herbaceous. Peduncle spreading, 1-2 cm long, obcompressed. Umbels of both sexes 10- to 25-flowered, 1-2 cm in diam. at anthesis; rays 5-10 mm long, slender. Staminate perianth segments 5-6 mm long, reflexed at anthesis; outer ones oblong 0.9-1.2 mm wide, obtuse, inner ones linear to linear-oblanceolate, 0.3 mm wide. Stamens 6; filaments 4-4.5 mm long; anther oblong, 1 mm long, white. Pistillate perianth segments 4-5 mm long, recurved at anthesis, outer ones lanceolate-ovate, 1-1.2 mm wide, subobtuse, inner ones linear 0.2-0.3 mm wide, obtuse. Ovary ellipsoid, 1.5 mm long, 1 mm wide; stigmas 0.8-1 mm long, suberect; staminodia 3, needle-like, 1-1.5 mm long.
• Berries ellipsoid, 5-7 mm across, dirty yellow-brown then turning purple-brown at maturity. Seeds 1-3, orbicular, thickly biconvex, ca. 3 mm long, dark red.

物候

Flowering in winter.

分布

• 台灣 : occasional in forests, 100-1,800 m.
• 世界 : Indochina and the Philippines through Taiwan and the Ryukyus northward to Kyushu. Evergreen forests at relatively high elevations, in Japan descending to sea level.

引證標本

TAIPEI: Hsiaoketou, Wang 762; Tatunshan, T. Suzuki 6773; Tungyenshan, Lin & Liu 96. ILAN: Shenmihu, Huang & Huang 13393; Wushihpi, Chen 80. TAOYUAN: Paling, Peng 11043. HSINCHU: Hsinpu, Wang 757. TAICHUNG: Toukeshan, Peng 9962. NANTOU: Lienhuachih, Chiu et al. 2829; Pilu, Wang 1312. CHIAYI: Alishan, Chen 10684 Taiping, Lin & Chen 509. KAOHSIUNG: Kuaiku, Tzeng 276; Shanping, Chang 7024. PINGTUNG: Chiupeng, Y. Tateishi et al. s. n.; Laofoushan, Chen et. al. 1024; Sheting, Chang 3400; TAITUNG: Chachayalaishan, Yang et al. 25001 Lanyuh, Chang 2820; Lutao, Chang 16572. HUALIEN: Hoping, Hsieh et al. 532; Shenmiku, Chen & Lien 322.

參考文獻

• KOYAMA, Tetsuo, WANG, Kuang-Yuh, LU, Fu-Yuan, and OU, Chern-Hsiung, Flora of Taiwan, second edition 5: 103. 2000.  (Fl. Taiwan 2nd edit.)