生活型

liana

形態特徵

• Climbing vines. Stems terete, smooth, 1-5 m long, 3-5 mm thick; branches; weakly to hardly zigzag, internodes terete, smooth, 1-5 cm long, 1-2 mm thick.
• Leaf blade lanceolate-oblong to lanceolate-elliptic or ovate, 5-10(-15) cm long, 2-3.5(-5) cm wide, base suddenly contracted or rounded, apex gradually or abruptly attenuate to acute or acuminate, coriaceous, lower surface pale or white-powdery; costas 5 including a marginal pair, all separate to base, raised on lower surface, intercostal areas on both sides of midvein much broader than outer areas, lateral veinlets oblique to divergent, conspicuous, forming dense reticulations; petiole 1-3 cm long, 1-1.2 mm thick, laterally compressed, breaking off at apex, sheathing for 3-15 mm long; wings 0.3-1 mm wide, herbaceous, not auriculate; tendrils slender, 7-15 cm long.
• Umbels in non prophyllate leaf axils; peduncle 1-9 mm long; bracteoles ovate-acute, 1-1.7 mm long. Staminate umbels globose, flowers 30-60, rays capillary, 1-2.5 cm long; pistillate umbels turbinate to hemisphaerical, flowers 10-25, rays filiform, 1-1.5 cm long. Staminate perianth depressed globose, 2 mm long, 3-3.7 mm across, obtusely trigonous, pale greenish, tepals free, only sligtly open; outer tepals saccate, deeply furrowed along median line, transversely widely obovate-orbicular, 3 mm wide; inner tepals ovate, acute, 1.5 mm long, 1 mm wide, boat-shaped, dorsally furrowed, strongly incurved. Stamens 6; anthers 0.7 mm long, white, subsessile. Pistillate perianth pale. depressed globose, 1.5-2 mm long. 2.5-3 mm across; tepals free, incurved, outer ones orbicular or obovate-orbicular, 2 mm wide, rounded, boat shaped, inner ones obovate, shallowly saccate, 1.5 mm wide. Ovary ellipsoid, 3-furrowed, ca. 2 mm long and wide; stigmas annulate, 3-lobed; staminodia 3, needle-like, 2 mm long.
• Berries globose, 5-8 mm in diam., blue-black; seeds 1-3.

物候

flowering in summer and autumn.

分布

• 台灣 : occasional in forests, 300-1,000 m.
• 世界 : India, Myanmar, Thailand, Indochina, central and southern mainland China.

引證標本

TAIPEI: Hsiaoketou, Huang 394; Peitou, Janaka 29852. TAOYUAN: Taoyuan, Liu 778. HSINCHU: Chienshih, Wang & Lin 1975. TAICHUNG: Pahsienshan, Liu et al. 478; Takeng, Wang 1195. NANTOU: Huisunglinchang, Ou et al. (S5116); Kuantaohsi, Hsieh 1451; Lienhuachih, Y. Yamamoto (29865, 30178); Lienhuachih, Wang 1212; Sun Moon Lake, Kudo & Sasaki 15273.

參考文獻

• KOYAMA, Tetsuo, WANG, Kuang-Yuh, LU, Fu-Yuan, and OU, Chern-Hsiung, Flora of Taiwan, second edition 5: 114. 2000.  (Fl. Taiwan 2nd edit.)