特徵描述
作者
SU, Horng-Jye
型態特徵
Terrestrial, epiphytic or saprophytic perennial herbs with rhizomes, corms, root-stem tuberoids or tuberous roots. Stems monopodial or sympodial, leafy or scapose, long or short, often thickened and forming pseudobulbs. Leaves radical or cauline, spiral or alternate and often distichous, convolute or conduplicate, variously shaped, sometimes all reduced to scales, often sheathed at base, sheath nearly always closed and encircling stem. Inflorescences terminal or lateral, spicate, racemose or paniculate, or with solitary flower; peduncles from various positious of stem, scapes from rhizome or base of pseudobulb. Flowers usually zygomorphic, often resupinate through twist of ovary for 180o, perianth epigynous, composed of 6 petaloid segments in 2 whorls, free or variously connate in each whorl; sepals (3 outer segments) imbricate or subvalvate; petals (3 inner segments) usually alternate to sepals, middle petal (lip or labellum) usually different in shape and often complicated in structure; lip variously shaped, sometimes 3-lobed, mostly placed in an abaxial position on account of resupination, base often produced into sac or spur; stamens and pistil fused together forming column, rarely separated; stamens 1 or 2, anthers 2-8-locular, sometimes with reduced partitions, opening by lengthwise slit, pollens rarely granular, generally agglutinated into masses or compact bodies (pollinia); pollinia mealy (powdery), sectile (consisting of small massulas) or waxy (bony), 2, 4, 6 or 8, naked (without definite accessory structure), or with accessory structures forming set of pollinarium, either with one end extended into softer sterile caudicle developed in anther and attached by caudicle to viscidium, or connected by sterile hyaline stipe developed outside anther, stipe derived from epidermis of rostellum (tegula stipe) or formed by apex of rostellum (hamulus stipe); ovary inferior, usually 1-locular with 3 parietal placentas, often produced at apex into semiterete column bearing anther and stigma; stigma basically 3-lobed or with 3 stigmatic areas, lobes usually confluent in single concave or convex area on ventral side of column, or reduced to 2 discrete lateral lobes on ventral or lateral sides of column, apex of median lobe often transformed into small outgrowth (rostellum) lying between anther and receptive stigmatic areas, portion of rostellum sometimes modified into small viscid disk or viscidium attaching to pollinia or their stipes; ovules numerous and minute, anatropous. Fruits usually capsule, mostly opening laterally by 3 or 6 longitudinal slits. Seeds numerous, usually minute, dust-like, without endosperm; embryo not differentiated.
屬
About 500 genera and more than 24,000 species, cosmopolitan, mainly in tropical and subtropical regions, extending into temperate and subarctic areas; 101 genera with 322 species, 3 subspecies and 5 varieties in Taiwan.
參考文獻
- SU, Horng-Jye, Flora of Taiwan, second edition 5: 729. 2000. (Fl. Taiwan 2nd edit.)
- Dressler, R. L., Dioscorides Press, Oregon. 314pp. 1993.
Gastrodia 赤箭屬
作者
LEOU, Chong-Sheng
型態特徵
Terrestrial saprophytic herbs without chlorophylls. Rhizomes horizontal, tuberous, densely noded, often pubescent, with scales on node, rarely with coral-like roots. Stem erect, scaly on nodes. Leaves absent. Inflorescence terminal, racemose, bearing few to many bracteate flowers. Flowers resupinate, fleshy; ovary pedicellate, pedicels elongated in fruit or not; sepals and petals united, forming a 5-lobed perianth tube, or forming a urceolate structure being split between lateral sepals, petal lobes located near sinus of sepal lobes, smaller than sepal lobes; lip adnate to apex of column-foot, embraced by perianth tube, or rarely adnate to perianth tube, not lobed or rarely 3-lobed, clawed at base, blade often mobile, disc with or without lamellae or calli; column fairly long, erect on top of ovary, more or less semicylindrical, straight, narrowly 2-winged near apex, often with foot at base, rarely with ventral appendage; anther terminal on column, incumbent, pollinia 2, soft, granular, without viscidium; stigma single, near base or rarely near top on ventral side of column; rostellum small or rarely absent. Capsules cylindrical or ellipsoid.
參考文獻
種列表
- Gastrodia albida 白赤箭
- Gastrodia appendiculata 無蕊喙赤箭
- Gastrodia callosa 緋赤箭
- Gastrodia clausa 閉花赤箭
- Gastrodia confusa 八代赤箭
- Gastrodia confusoides 擬八代赤箭
- Gastrodia elata 高赤箭
- Gastrodia flavilabella 夏赤箭
- Gastrodia flexistyla 摺柱赤箭
- Gastrodia fontinalis 春赤箭
- Gastrodia gracilis 山赤箭
- Gastrodia kaohsiungensis 高雄赤箭
- Gastrodia nantoensis 南投赤箭
- Gastrodia nipponica 日本赤箭
- Gastrodia peichatieniana 北插天山赤箭
- Gastrodia pubilabiata 冬赤箭
- Gastrodia rubinea 紅寶石赤箭
- Gastrodia shimizuana 清水氏赤箭
- Gastrodia sp.
- Gastrodia stapfii 黃赤箭
- Gastrodia sui 蘇氏赤箭
- Gastrodia theana 短柱赤箭
- Gastrodia uraiensis 烏來赤箭