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Gramineae

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特徵描述
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特徵描述

作者

HSU, Chien-Chang

型態特徵

Annuals or perennials herbs, rather rarely with woody perennials culms (subfamily Bambusoideae); hollow or solid, septate at nodes. Perennials herbs or their branches at base usually with scales (cataphylls, transformed leaf sheaths). Leaves two-ranked, consisting of a sheath, ligule, and a blade. Axillary buds when grown out into a branch provided with a usually 2-nerved bract (prophyll) between branch and culm. Leaf blade mostly linear and not petiolate at base, rarely with broad blades and stipitate (pseudo-petiole) at base (especially in Bambusoideae); ligules membranaceous or ciliate, rarely lacking (Echinochloa), margins of sheath of leaf connate at least at base. Inflorescences usually terminal, primarily paniculate, but secondarily extremely variable and transformed into racemes, spikes, digitate umbels, etc. Flowering unit (spikelet) usually with 2 bracteoles (glumes) at base; base occasionally with an obconical blunt to very acute thickening (callus). Flowers alternatingly along a central axis (rachilla) supported by a bract (lemma) and a prophyll (palea), bisexual or occasionally unisexual. Lemma and palea when falling as a unit clasping fruit at base often with an obconical blunt to very acute thickening (also called callus). Perianth usually composed of 2 or 3 minute scales (lodicules), rarely 6, sometimes absent (e.g. in sterile or female florets). Stamens usually 3, sometimes 6 (Bambusoideae), or 1 or 2; filaments filiform; anthers 2-locular. Pistil 1, superior; ovary and single anatropous ovule completely adnate; styles 2 or 3, apical or subapical, stigmas plumose. Caryopsis with rich endosperm; embryo basal, abaxial (facing lemma), opposite to punctiform to linear adaxial hilum (facing palea).

The Gramineae are one of the largest families of vascular plants with an estimated 620 genera and 10,000 species. Keng (1965) reported 201 genera and 775 species for mainland China. Hsu (1974) treated 5 subfamilies (excluding Bambusoideae), 27 tribes, 118 genera, 289 species, 1 subspecies, 39 varieties, and 8 forms in Taiwan.

The following arrangement of subfamilies, tribes and genera is that of Hsu (1978) with very few modifications. The descriptions of most genera are unchanged in this edition except that genera of the subfamily Pooideae (Festucoideae) which was mostly adopted from Clayton and Renvoize (1986). Some critical remarks were provided by Dr. J. F. Veldkamp, National Herbarium of the Netherlands, Leiden. Some line-drawings of Paniceae were drawed by Y. Fujimoto, Botanical Institute of Fujimoto, Kobe, Japan.

參考文獻

Leptaspis 囊稃竹屬

作者

HSU, Chien-Chang

型態特徵

Monoecious perennials; culms solid, erect or ascending. Leaves strikingly distichous; sheaths laterally compressed; ligule very short; blade with a distinct pseudo-petiole, with abaxial side upwards, usually lanceolate, pinnate nerved, with numerous distinct transverse nerved. Inflorescence umbel-like or paniculate, with branches borne on all sides. Spikelets unisexual, 1-flowered, ultimate branchlets with a small male spikelet at top, with 1 or more larger female spikelets below. Male spikelets: glumes thinly membranaceous; lemma larger, conduplicate with free margins, 5-9-nerved; palea entire or shortly bidentate, 2-nerved; lodicules wanting; stamens 6, erect, filaments short, anthers linear, only tips protruding. Rudiment of pistil obsolete or minute; spikelets soon deciduous. Female spikelets: glumes thinly membranaceous; lemma much larger and firmer, usually pear-like, inflated, margins connate, leaving a small, terminal or lateral pore, 5-9-nerved, with short hooked hairs all over, enlarged and usually indurated at maturity; palea narrow, free or adnate to margins of lemma, grooved whole length on hilum-side.

參考文獻

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