特徵描述
作者
STAPLES, George William and YANG, Sheng-Zehn
型態特徵
Herbs or shrubs, stems usually twining or scandent, less often prostrate or erect, sap often milky. Leaves alternate, simple, entire, dissected, or compound (scales in Cuscuta). Flowers solitary, axillary or in cymes, racemes, panicles, umbels, or capitula, hermaphroditic, actinomorphic, usually 5-merous, often showy. Sepals free, often persistent, sometimes accrescent in fruit. Corolla sympetalous, funnelform, campanulate, salverform, or urceolate; limb nearly entire or deeply lobed. Stamens alternating with corolla lobes, adnate to corolla; filaments filiform, equal or unequal in length; anthers introrse, laterally and longitudinally dehiscing; pollen smooth or finely spiny. Disc ring-like or cupular. Ovary superior, mostly 2-carpellate, 1- or 2-loculed, rarely 3- or 4-loculed; ovules basal, erect; style(s) 1 or 2, terminal (gynobasic in Dichondra) or very short or absent; stigma entire or lobed, rarely peltate. Fruit a capsule, dehiscing by valves, circumscissile, or irregularly shattering, less often a berry or nut-like. Seeds usually trigonous, smooth or pubescent.
Pollen aperture type and surface ornamentation are important characters in classification of Convolvulaceae at generic level and above. The most critical feature of the pollen is whether the grain surface is spiny or not. A 20 × hand lens or the low power of a dissecting microscope is adequate for discerning the presence or absence of minute spines on the surface. The number and arrangement of apertures is of secondary importance.
For successful identification of Convolvulaceae, both flowers and fruits are necessary. The following key to genera assumes that adequate fertile material with both flowers and fruit is available; sterile collections are hopeless unless the user of the key is well acquainted with the taxa in their fertile condition. Furthermore, the key requires use of a pollen character at a single point in order to proceed, although pollen characters are mentioned elsewhere as confirmatory evidence.
屬
With 58 genera and an estimated 2,000 species, Convolvulaceae are widely distributed in tropical, subtropical, and temperate regions. There are 14 genera and at least 44 species in Taiwan. Several additional taxa that are widespread weeds will likely be found in Taiwan in the future.
參考文獻
- STAPLES, George William and YANG, Sheng-Zehn, Flora of Taiwan, second edition 4: 341. 1998. (Fl. Taiwan 2nd edit.)
- Austin, D. F., D. A. Powell & D. H. Nicolson., Brittonia 30(2): 195-198. 1978.
- Chang, C. E., Flora of Taiwan 4: 347-389. 1978. (Fl. Taiwan)
- Fang, R. C. & G. W. Staples., Flora of China Family 16: 271-325. 1995. (Fl. China)
- Fang, R. C. & Huang S. H., Flora Reipublicae Popularis Sinicae 64(1): 11-53. 1979. (Fl. Reip. Pop. Sin.)
- Ferguson, I. K., B. Verdcourt & M. M. Poole., Kew bulletin 31(4): 763-773. 1977. (Kew Bull.)
- Lu, F. Y., Quarterly journal of Chinese forestry 6: 105-165. 1972. (Quart. J. Chin. For.)
- Nicolson, D. H., C. R. Suresh, & K. S. Manilal, An interpretation of Van Rheedes Hortus Malabricus Regnum Veg. 119: 1-378. 1988.
- Ooststroom, S. J. van, Flora Malesiana : Series I : Spermatophyta 4(4): 388-512. 1953. (Fl. Males. Ser. I)
Merremia 菜欒藤屬
作者
STAPLES, George William and YANG, Sheng-Zehn
型態特徵
Herbs or shrubs, often twining, sometimes prostrate. Leaves usually petiolate, rarely sessile, margin entire or dentate, lobed, or palmately or pedately lobed or compound. Flowers axillary, solitary or in few- to many-flowered variously branched cymose inflorescences; bracts usually small. Sepals variable in shape, often convex, nearly equal or outer 2 sepals smaller, persistent, often enlarged in fruit. Corolla often yellow or white, sometimes with a darker center, funnelform or campanulate, usually glabrous, or midpetaline bands + sericeous, sometimes only at apex; limb entire or + 5-angled. Stamens included, often unequal; anthers often spirally twisted; filaments dilated basally, filiform distally; pollen 3–12-colpate or polyrugate, not spiny. Disc ring-like. Pistil included; ovary (imperfectly 2) 4-loculed, 4-ovuled; style 1, filiform; stigmas 2, globular. Capsule 1–4-loculed, usually 4-valved or + irregularly dehiscing. Seeds 4 or fewer, glabrous or pubescent to villous especially at margin.
Merremia is often confused with Ipomoea, which has pantoporate, spiny pollen, straight anthers and very few species with yellow corollas, and Operculina, which consistently has fruits with a circumscissile lid that detaches first, leaving a fragile endocarp that shatters irregularly. The distribution of hairs on the corolla is taxonomically important in Merremia; this is most easily discerned on mature flower buds.