生活型

herb

形態特徵

• Annual herb, up to 140 cm tall. Stems erect or ascending, simple or branched from base, glabrous to pilose, basal internodes 0.6–2 cm long, upper internode to 9 cm long.
• Lowermost leaf blade broad ovate, subcordate, with attenuate petioles, margins strongly dentate, ca. 4 cm long, 4.5 cm broad, petioles 1.5–2.5 cm long, moderately pilose, wingless and not auriculate; lower (2nd and 3rd from the bottom) leaf blades broad ovate, subcordate, with attenuate petioles, leaf margin broad dentate, ca. 6 cm long, 6 cm wide, petioles 2–3 cm long, winged, not auriculate, moderately pilose; middle part leaves similar to the lower ones but strongly auriculate, auricles broadly dentate; uppermost leaves sessile, auriculate-cordate, more or less deltoid, midrib of the lower surface moderately pilose.
• Capitula erect, discoid, solitary to 7 in open corymbs; bracts narrowly lanceolate, apex attenuate, 8–11 mm long, pilose; capitula in anthesis ca. 20 mm long, 12 mm in diameter; involucre tubular, ca, 10 mm long, base 4 mm wide, middle part 3 mm wide, phyllaries uniseriate, 9–12, moderately pilose; florets ca. 80, tubular, 5-lobed, yellowish, orangey or cream white, corolla lobes purple or orange-tinged, corollas ca. 8 mm long, lobes 2 mm long. Anthers dark orange, 1.7 mm long, apex appendiculate. Style branches orange, ca. 1.2 mm long, recurved. Pappus ca. 7 mm long, finely barbellulate.
• Achenes truncate-elliptic, 3 mm long, 5-ribbed, hispid along achene midribs.
• Chromosome number, 2n = 20.

物候

flowering and fruiting all year round.

引證標本

Keelung City: Hsishih Dam, ca. 50 m, 31 Jan 1997, C.-I Peng et al. 16938 (HAST), same locality, 65 m, 4 Mar 2004, C.-I Huang et al. 1628 (HAST); Hopingdao, 16 Dec 2003, M.-J. Jung s.n. (TAIF); Hsinshan Dam, 50–150 m, 16 Apr 2005, J.-H. Lii & S.-W. Chung 1174 (TAI); Anle District, Chingjenhu, 187 m, 16 Mar 2007, T. Y. A. Yang et al. 19245 (TNS). Ilan County: Chuangwei Township, Kuoling, 24 Apr 1999, K.-F. Chung et al. 1153 (HAST), Yungchen village, ca. 0 m, 14 Jun 1999, C.-I Peng et al. 17251 (HAST), near Lanyang Bridge, 5 m, 4 Aug 2004, C.-I Huang et al. 1796 (HAST); Yuanshan Township, 300–400 m, 17 Jan 2005, S.-W. Chung 8288 (TAIF). Taipei County: Hsichih City, ca. 230, 2 Jan 2000, C.-I Peng 17925 (HAST), 1 Jan 2008, M.-J. Jung 2117 (TAIF), Chiangtzu Gazebo, 530 m, 6 Mar 2000, C.-H. Lin & K.-F. Chung 76 (HAST), same locality, 3 Aug 2001, S.-C. Wu et al. 2449 (HAST, TAI); Juifang Township, Houtung, 150 m, 8 Sep 2001, K.-Y. Wang & K.-H. Chen 1654 (TAIF), Shumeiping, 630 m, 15 Mar 2003, T. Y. A. Yang et al. 15618 (TNS); Pingsi Township, 14 Nov 2007, M.-J. Jung 1877 (TAIF), Shihfen, 3 Apr 2008, M.-J. Jung 2649 (TAIF), Siaozihshan, 300–360 m, 2 Mar 2003, S.-C. Liu & P.-C. Liao 990 (HAST); Shuangsi, Dapishan, 400–515 m, 23 May 2003, S.-C. Liu et al. 1196 (HAST, TAIF, TNS), Sanchakang Village, 100 m, 4 Oct 2008, P.-F. Lu 17002 (TAIF). Taipei City: Beitou, 160 m, Chunchienyen, 15 Mar 2007, S.-S. Dai 1948 (TAIF); Daan, 20 m, Guting, 5 Apr 2008, M.-J. Jung 2657 (TAIF); Nankang, Academia Sinica, 30 m, 14 Dec 2002, S.-M. Ku 1688 (HAST); Xinyi, Hsiangshan, 200 m, 28 Mar 1999, W. C. Leong 1148 (HAST, TNS); Zhongshan, Tachia River Park, 15 m, 21 Mar 2007, C.-I Huang & K.-F. Chung 3096 (HAST). Miaoli County: Sanyi Township, Shengsing Railway Station, 380 m, 21 Jul 2008, C.-Y. Lin et al. 13 (HAST).

引用文獻

• Kuo-Fang Chung, Shin-Ming Ku, Yoshiko Kono and Ching-I Peng, Taiwania 54(4): 385-390. 2009.