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Plants of Taiwan

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    台灣植物誌 第二版  Flora of Taiwan, 2nd edition    Vol. 3

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    台灣植物誌第二版  Flora of Taiwan, 2nd edition  3: 674

     78. ICACINACEAE 茶茱萸科
    CHANG, CHING-EN

    Trees and shrubs, often climbing. Leaves usually alternate, simple, estipulate. Flowers bisexual, rarely unisexual by abortion, actinomorphic, usually in panicles. Calyx 4-5-lobed, small. Petals 4-5, usually distinct or sometimes connate, valvate. Stamens as many as petals and alternate with them, distinct, the anthers 2-celled, usually introrse, sometimes 4-lobed, dehiscing longitudinally. Pistil 1, the ovary superior, 1-loculed, mostly 3- or 5-carpellate, the placentation apical; ovules 2, usually pendulous from locule apex, anatropous; style 1, the stigmas usually 3. Fruit usually a drupe, rarely a samara. Seed with straight or curved embryo; endosperm present.
    About 56 genera, of pantropical distribution. Three in Taiwan.

    KEY TO GENERA
    1. Calyx cupular, minutely 4-5 dentate, valvate in bud.
    2. Inflorescences terminal; filaments glabrous… 3. Nothapodytes
    2. Inflorescences axillary; filaments with subclavate hairs… 1.Gomphandra
    1. Calyx deeply 5-parted, imbricate in bud… 2. Gonocaryum

    1. GOMPHANDRA Wall. ex Lindl. 毛蕊木屬

    Dioecious trees or shrubs. Branchlets slender, smooth. Leaves spiral, chartaceous to coriaceous, often tubercled beneath, penninerved. Flowers in short cymes, these several from an axillary peduncle, the male many, the female few flowered; calyx cupular, minutely 4-5-toothed; petals 4-5, free or coherent below. Disk absent or obscure. Male flowers: stamens 4-5, usually exserted; filaments flattened, with subclavate hairs on the ventral side below the anthers and dorsally at the connective; rudiment of ovary conspicuous. Female flowers: petals sometimes absent; ovary cylindric; stigma sessile, discoid, slightly 4-5-lobed or 4-5 crenate. Drupes ellipsoid or oblong, sometimes shortly beaked, base sometimes swollen; endocarp crustaceous or hard, with 8-10 longitudinal ridges especially prominent in the dry fruit.
    About 33 species in SE. Asia(India, Ceylon to Burma, Thailand, Indo-China, and S. China including Hainan) and in Malaysia. One species in Taiwan.

    1. Gomphandra luzoniensis (Merr.) Merr., Enum. Philip. Fl. Pl. 2: 490. 1923. 呂宋毛蕊木 Pl.349
    Stemonurus luzoniensis (Merr.) Howard in Journ. Arnold Arb. 21: 468. 1940; Ou in Quart. Journ. Chin. For. 9(2): 119. t. 2. 1976.
    A small tree. Branchlets glabrous, terete. Leaves obovate-elliptic, obtuse at apex, obtuse-rounded at base, glabrous, 7-9 cm long, 5-6.5 cm broad, the lateral veins 3-4 on per side, veinlets obsolete; petioles stout, strongly grooved above, 1-1.5 cm long, glabrescent. Inflorescences axillary with 3-5 cymes on a common peduncles, densely pubescent; calyx cupular, minutely 5-toothed, glabrescent; petals 5, whitish, 4-5 mm long, glabrous, oblong; stamens 5, the filaments flattened, with long hairs below the anthers and on the connective; ovary cylindric, glabrous; stigma discord; style none. Drupe oblong, about 2 cm long, 0.8-1 cm across, glabrous, with many longitudinal ridges.
    The Philippines. Taiwan, on Is. Lanyu and Is. Lutao.