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Plants of Taiwan

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    台灣植物誌 第二版  Flora of Taiwan, 2nd edition    Vol. 3

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    台灣植物誌第二版  Flora of Taiwan, 2nd edition  3: 661

     76. STAPHYLEACEAE 省沽油科
    LI, HUI-LIN

    Trees and shrubs. Leaves opposite, pinnate, rarely 1-foliolate or ternate, stipulate; leaflets serrate. Flowers small, polygamo-dioecious, in terminal or axillary clusters. Calyx mostly 5-lobed, imbricate. Petals 5, free, imbricated in bud. Stamens 5, alternate with petals, inserted outside the cup-shaped disc. Ovary usually 3-celled, entire or lobed, the styles connate or distinct, terminal. Ovules 6 or numerous in each cell; placentation axile. Fruit a leathery or fleshy capsule, sometimes indehiscent. Seeds solitary or few in each cell, albuminous.
    About five genera and 25 species, of wide distribution; two in Taiwan.

    KEY TO GENERA
    1. Leaves pinnate, the leaflets 5-7; fruit of 1-3 coriaceous follicles; seeds arillate… 1. Euscaphis
    1. Leaves 1- or 3-foliolate; fruit berry-like, globose; seeds not arillate… 2. Turpinia

    1. EUSCAPHIS Sieb. & Zucc. 野鴉椿屬

    Small tree, glabrous and shining. Leaves opposite, odd-pinnate, stipulate; leaflets opposite, serrulate, chartaceous. Flowers small, yellowish white, in terminal panicles; calyx lobes 5, imbricate; petals 5, imbricate; stamens 5, inserted outside the cup-like disc, the filaments short, the anthers sub-rounded; ovary 2-3-lobed and celled; styles 2-3, combined or connivent, the stigmas subcapitate; ovules several in each cell, in two rows. Fruit of 1-3 follicles, the pericarp coriaceous, dehiscent along the ventral sutures; seeds l-3 in each follicle, arillate.
    Monotypic, E. Asia.

    1. Euscaphis japonica (Thunb.) Kanitz in Termeszet. Fuzet. 3: 157. 1878; Li in Fl. Taiwan 3: 636. 1977. 野鴉椿 Pl.342, Photo 49
    Sambucus japonica Thunb., Fl. Jap. 125. 1784.
    A deciduous shrub or small tree, glabrous. Leaves opposite, 5-7-pinnate, the leaflets opposite, ovate to ovate-lanceolate, 5-8 cm long, 3-4 cm broad, acute to acuminate, rounded to obtuse or broadly cuneate at base, appressed serrulate, glabrous above, glabrous beneath or villose along the veins toward the base, the rachis reddish. Inflorescence a broad many-flowered long-peduncled panicle, 5-12 cm across; flowers small, pentamerous; sepals, petals, and stamens of nearly equal length of about 2 mm; ovary 2-3-lobed, about 2 mm long, glabrous; pedicels 2-4 mm long, glabrous. Fruit of 1-3 spreading leathery dehiscent pods, subglobose to falcate-ovoid, about 1.2 cm long, slightly reddish.
    Southeastern to central China, Hainan, the Ryukyus to Japan. Taiwan, in forests, mountains of northern Taiwan.
    TAIPEI: Kudo, Suzuki & Mori 330; Kao s. n. 1957*.

    2. TURPINIA Ventenat 山香圓屬

    Trees or shrubs, glabrous and shining, with smooth terete branches. Leaves opposite, stipulate, 1- or 3-foliolate to odd-pinnate; leaflets opposite, serrulate, subcoriaceous. Panicles terminal and axillary. Flowers small, regular, hermaphroditic; calyx 5-partite, imbricate; petals 5, imbricate; stamens 5, inserted