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Plants of Taiwan

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    台灣植物誌 第二版  Flora of Taiwan, 2nd edition    Vol. 3

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    台灣植物誌第二版  Flora of Taiwan, 2nd edition  3: 510

     63. RUTACEAE 芸香科
    CHANG, CHING-EN and HARTLEY, TOM G.
    Acronychia, Melicope, Tetradium by HARTLEY, TOM G.

    Trees, shrubs, rarely herbs, with abundant pellucid glands, filled with essential oils. Leaves opposite or alternate, simple or compound, estipulate. Inflorescence various. Flowers usually regular, hermaphroditic or polygamous; sepals 4-5, free or connate; petals 4-5, rarely none, imbricate or valvate; stamens as many or twice as many as the petals, sometimes fewer or more numerous; disc usually annular or cushion-like, between the ovary and stamens; ovary superior, usually deeply lobed, typically 4-5-loculed, the carpels more or less connate, or free, rarely 1-carpellary; ovules solitary or geminate. Fruit various, separating into as many cocci as carpels or baccate. Seeds usually solitary in each cell, the testa various; endosperm fleshy or 0; embryo straight or curved.
    About 140 genera with 1,300 species, tropical and temperate; 13 genera in Taiwan.

    KEY TO GENERA
    1. Gynoecium 1- to 5-carpellate, the carpels free or connate up to about 1/2 their length; fruit dehiscent.
    2. Plants ligneous; carpels with 2 ovules.
    3. Leaves alternate… 13. Zanthoxylum
    3. Leaves opposite.
    4. Inflorescences terminal or terminal and upper axillary; leaves imparipinnate… 11. Tetradium
    4. Inflorescences axillary and/or on branchlets below leaves; leaves digitately 3-foliolate or 1-foliolate… 6. Melicope
    2. Plants herbaceous; carpels usually with 6-8 ovules… 2. Boenninghausenia
    1. Gynoecium 2- to 18-carpellate, the carpels connate for their full length (rudimentary carpels free or not in male flowers of Phellodendron); fruit indehiscent.
    5. Stamens at most twice the number of petals.
    6. Leaves simple or 1-foliolate.
    7. Leaves alternate.
    8. Stem armed with prickles… 9. Severinia
    8. Stem unarmed… 10. Skimmia
    7. Leaves opposite… 1. Acronychia
    6. Leaves compound; leaflets 3 or more.
    9. Stem armed with prickles… 12. Toddalia
    9. Stem unarmed
    10. Leaves alternate
    11. Styles persistent; overy with only 1 pendent ovule in each locule… 5. Glycosmis
    11. Style deciduous: ovary with 2 ovules in each locule.
    12. Flower buds globose or broadly ovoid; styles short and thick, the stigma flattened, scarcely broader than style… 4. Clausena
    12. Flowers buds cylindrical or narrowly ovoid; style long and slender, usually longer than the ovary, the stigma broader than the style… 7. Murrava
    10. Leaves opposite… 8. Phellodendron
    5. Stamens more than twice the number of petals… 3. Citrus

    1. ACRONYCHIA J. R. & G. Forst. 降真香屬
    HARTLEY, TOM G.