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Plants of Taiwan

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    台灣植物誌 第二版  Flora of Taiwan, 2nd edition    Vol. 1

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    台灣植物誌第二版  Flora of Taiwan, 2nd edition  1: 99

     11. HYMENOPHYLLACEAE 膜蕨科
    TSAI, JINN-LAI and SHIEH, WANG-CHUENG

    Mostly small epiphytic, petrophilous or terrestrial, usually delicate plants; rhizomes usually longcreeping and slender, covered with hairs, bearing distant fronds, or sometimes short and erect with clustered fronds. Fronds usually small, of many shapes; the blades only 1-cell thick, simple or variously divided, the veins free, the ultimate segments with only a single vein. Sori marginal or extramarginal, on a slender extension of the vein constituting a receptacle, surrounded by a tubular or sometimes 2-valved involucre-like indusium; sporangia borne on an elongate or capitate receptacle, sessile, the annulus complete, oblique, the dehiscence vertical. Spores trilete. Gametophytes filamentous (algal-like) or ribbon-like.
    A family of about 34 genera and 700 species, mostly tropical and subtropical, 15 genera in Taiwan.

    KEY TO GENERA
    1. Rhizomes short and erect with tufted fronds.
    2. Small epiphytes; laminar cells transversely elongate… 1. Abrodictyum
    2. Terrestrial ferns; laminar cells not transversely elongate.
    3. Fronds pinnate; indusia truncate… 3. Cephalomanes
    3. Fronds bipinnatifid to 4-pinnatifid; indusia not truncate.
    4. Laminae bipinnatifid to tripinnatifid; indusia 2-lipped and notched… 14. Selenodesmium
    4. Laminae 4-pinnatifid; indusia tubular.
    5. Stipes and rachis with long multicellular hairs; indusia entire… 2. Callstopteris
    5. Stipes and rachis without long multicellular hairs; indusia with clavate hairs on their margins… 11. Nesopteris
    1. Rhizomes long-creeping and slender with distant fronds.
    6. Indusia 2-lipped, cleft to base or nearly so.
    7. Margins of fronds and indusia entire… 7. Mecodium
    7. Margins of fronds and indusia toothed.
    8. Indusia cleft nearly to base; receptacles included… 6. Hymenophyllum
    8. Indusia tubular towards base; receptacles long exserted… 8. Meringium
    6. Indusia tubular or conical.
    9. Laminae with false veins, either occuring in a submarginal line or scattered short striae.
    10. Fronds simple, rarely lobed… 9. Microgonium
    10. Fromds digitately or pinnately divided to tripinnatifid.
    11. Marginal cells differing from laminar cells… 13. Reediella
    11. Marginal cells not specialized… 4. Crepidomanes
    9. Laminae without false veins.
    12. Fronds digitatley and dichotomously lofed.
    13. Laminae fan-shaped, the upper margin deeply cleft but not dissected to base; indusia tubular, the mouth much enlarged… 5. Gonocormus
    13. Laminae not fan-shaped, dissected to base; indusia cone-like, the mouth deeply undulate or somewhat 2-lipped… 10. Microtrichomanes
    12. Fronds pinnatifid to 4-pinnatifid.
    14. Laminae covered with long brown hairs… 12. Pleuromanes
    14. Laminae without long hairs… 15. Vandenboschia