台灣植物誌 第二版 Flora of Taiwan, 2nd edition Vol. 1
5. PODOCARPACEAE 羅漢松科
LI, HUI-LIN and KENG, HSUAN
Podocarpus by YANG, YUEN-PO and LU, SHENG-YOUResinous trees or shrubs. Leaves alternate, rarely opposite, persistent, scale-like or needle-like, linear or lanceolate, sometimes broad and ovate, entire. Staminate and ovuliferous cones dioecious. Staminate cones axillary, the staminate scales with 2 pollen sacs. Ovuliferous cones consisting of a single (rarely 2) ovule enveloped by a layer of succulent structure ('epimatium') and seated on a receptacle formed by several fused scales which often becomes thickened or fleshy after fertilization. Seeds with a bony shell, surrounded by the fleshy epimatium; embryo with 2 cotyledons.
About 20 genera with over 120 species, mostly in southern hemisphere; the following two genera occur in Taiwan.
KEY TO GENERA
1. Leaves with many ± parallel veins; receptacle of the mature seeds not or only slightly swollened and hardly fleshy… 1. Nageia
1. Leaves with a single midrib; receptacle of the mature seeds swollen and fleshy… 2. Podocarpus
1. NAGEIA Gaertn. 竹柏屬
Columnar, evergreen trees. Leaves oblong-lanceolate to broadly ovate-elliptic, alternate or opposite, with many fine, nearly parallel lengthwise veins converging towards both ends. Staminate cones single or in clusters, axillary. Ovulate cone solitary or sometimes in pairs. Seeds drupe-like, usually globose, the receptacle as thick as the peduncle, sometimes slightly thickened and hardly fleshy.
About 5 species, from E. Himalayas to S. Japan and to Malesia; the following two species are found in Taiwan. This genus is formerly included in Podocarpus (sen. Lat.) as a section.
KEY TO SPECIES
1. Leaves 8-18 cm long; seeds 1.5-1.8 cm across… 1. N. fleuryi
1. Leaves 2.5-9 cm long; seeds 1.2-1.4 cm across… 2. N. nagi
1. Nageia fleuryi (Hickel) de Laub. In Blumea 32: 210. 1987. 長葉竹柏 Pl.216
Podocarpus fleuryi Hickel in Bull. Soc. Dendrol. France 57. 1930; H. Keng in J. Taiwan Mus. 40 (2): 30. f. 1-2. 1987.
Podocarpus nageia auct. Non R. Br.: Henry, List Pl. Form. 91. 1986. p. p.
Podocarpus wallichianus auct. Non Presl.: Yamamoto in J. Soc. Trop. Agric. 7: 147. f. 19. 1935.
Podocarpus sp. Li & H. Keng in Taiwania 5: 45. pl. 10. 1954.
Decussocarpus fleuryi (Hickel) de Laub. In J. Arnold Arb. 50: 355. 1969.
A tree. Leaves thick coriaceous, opposite and decussate, broadly lanceolate or narrowly elliptic, 8-18 cm long, 2-4.5 cm wide, the apex acuminate, the base gradually narrowly into a short petiole, 2-4 mm long. Staminate cones cylindric, 2-6 cm long, 3-6 in axillary cluster. Seeds globose, 1.5-1.8 cm across, bluish purple at maturity; peduncle scarred, about 2 cm long.