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    台灣植物誌 第二版  Flora of Taiwan, 2nd edition    Vol. 1

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    台灣植物誌第二版  Flora of Taiwan, 2nd edition  1: 264

     22. ONOCLEACEAE 球子蕨科
    DEVOL, CHARLES E. and SHIEH, WANG-CHUENG

    Rhizome creeping or erect, dictyostelic. Stipes not articulate to rhizome, covered with scales at base; fronds strongly dimorphic; sterile fronds herbaceous, pinnate to bipinnatifid; venation free or anastomosing; fertile fronds pinnate to bipinnate, dark to nearly black. Sporangia covered by an inner indusium and by coriaceous reflexed leaf margin. Spores green.
    Two genera, one in Taiwan.

    1. MATTEUCCIA Todaro 莢果蕨屬

    Large terrestrial ferns; rhizomes stout, erect or procumbent at base, dictyostelic; stipes clustered; scales on rhizome, stipe and rachis attached by base, thin, brown, lanceolate. Fronds dimorphic; sterile fronds bipinnatifid, texture herbaceous, the venation free, the undersurface of lamina sparsely covered with small narrow scales along costa and veins. Fertile fronds pinnate, much contracted, becoming purplish-brown, the margins coriaceous, inrolled, concealing the sori.
    Three species, one in Taiwan.

    1. Matteuccia orientalis (Hook.) Trev. In Atti 1st Veneto III. 14: 586. 1869; DeVol in Fl. Taiwan 1: 147. pl. 48. 1975. 東方莢果蕨 Pl.111
    Sterile fronds bipinnatifid; stipe 10-30 cm long, c. s. round, with 2 curved bundles; scales at base thin, brown, entire, lanceolate; lamina longer than wide; venation free, simple. Fertile fronds dark brown, persistent, pinnate; stipe 15-25 cm long; pinnae linear, usually 5-6 cm long. Sporangia in a coenosori, covered with thin, linear, inner indusium and coriaceous inrolled brown leaf margins.
    Distributed in Korea, Japan, Taiwan, China and Himalayas.
    ILAN: Taipingshan, Sasaki s. n. 1932*; Cross Island Highway, Ssuyuan, Hsu 853, Kuo 4040.